Apple - Migrate from HFS+ to APFS in-place
As of Jun 2017 according to this link, during macOS High Sierra install, file system can be upgraded from HFS+ to APFS:
How do I upgrade to Apple File System?
The macOS 10.13 installer offers nondestructive in-place upgrades from HFS+ to APFS for bootable volumes. You can use Disk Utility to convert external volumes from HFS+ to APFS format.
Note: this was true before the announcement of High Sierra, June '17
You cannot use APFS as a boot volume for Sierra, it's not yet supported.
APFS is still in development & will be capable of being used as a boot volume when it's finally released.
From Apple File System Guide - FAQ ...
Can I boot macOS Sierra from an APFS-formatted hard disk?
No. macOS Sierra supports Apple File System for data volumes only. You cannot boot macOS Sierra from a APFS-formatted volume.
How do I upgrade to Apple File System?
Apple will offer nondestructive in-place upgrades from HFS+ to APFS for all boot volumes when Apple File System ships in 2017. Tools will be available to convert external volumes from HFS+ to APFS format.