You uploaded a debuggable APK. For security reasons you need to disable debugging before it can be published in Google Play-Upload apk to google play

Don't use the debug variant output! Build a release apk. You can do that in Android Studio by going to the menu Build -> Generate Signed APK. Or by executing ./gradlew assembleRelease if you have properly configured signing in the build file.

I ran into this error and my application did not reference debuggable anywhere. After a little bit of searching, I found that I accidentally had testCoverageEnabled true in my release build type.

release {
    testCoverageEnabled true
    ...
}

Removing this resolved the issue.

I was having the same problem. Unknowingly I kept

debuggable true in release buildType

buildTypes {
        release {
            minifyEnabled false
            shrinkResources false
            debuggable true
            proguardFiles getDefaultProguardFile('proguard-android.txt'), 'proguard-rules.pro'

        }
    }

After changed to false. Its working fine.

buildTypes {
        release {
            minifyEnabled true
            shrinkResources true
            debuggable false
            proguardFiles getDefaultProguardFile('proguard-android.txt'), 'proguard-rules.pro'

        }
    }